Curriculum Vitae

1960-1963: Study of Biology/Chemistry at the Free University Berlin/Germany

1963-1967: Study of Biology/Genetics/Biochemistry at the Eberhard-Karls-University Tübingen/Germany

1967: Ph.D. at the Max-Planck-Institut für Biologie, Abt. Beermann, Tübingen/Germany

1967-1972: Research Associate, Max-Planck-Institut für Biologie, Abt. Beermann, Tübingen/Germany

1968: Postdoctoral Fellow, University of Edinburgh, MRC Group on the Mammalian Genome, Edinburgh/Scotland with P.M.B. Walker

1972: Habilitation “Biology”, Eberhard-Karls-University, Tübingen/Germany

1973: Habilitation “Molecular Biology”, University of Zürich/Switzerland

1973-1974: Oberassistent and Privatdozent, Institute of Molecular Biology II, University of Zürich/Switzerland

1974-1977: Head of an Independent Research Group at the Friedrich-Miescher-Laboratorium of the Max-Planck-Gesellschaft, Tübingen

1978-1997: Professor of Genetics, Head of Department of Genetics, Katholieke Universiteit Nijmegen/The Netherlands  (Emeritus since 1997)

since 1985: Teaching and Research in the Guest Laboratory of the Max-Planck-Gesellschaft, Institute of Cell Biology, Academia Sinica, 

since 1991 at the West China University of Medical Sciences, Chengdu/China

1991: Honorary Professor of the Academia Sinica, Institute of Cell Biology, Shanghai/China

1994: Honorary Professor of the West China University of Medical Sciences, Chengdu/China

1997- 2000: Guestprofessor, Faculty of Medicine, Johannes Gutenberg-University of Mainz/Germany

1998-2000: Member of the Managing Committee of the EBNIC (European Biotechnology Node for Interactions with China) an EU-funded activity of the EMBO (see:

1998: Honorary Professor (Faculty of  Biology), Johannes Gutenberg-University Mainz/Germany

2001: Honorary Professor Tongji Medical College Wuhan/China

2000-2008: Professor at the Shanghai Institutes for Biological Sciences (SIBS), Academia Sinica (long term lecturer of the German Academic Exchange Service (DAAD), and CAS-MPG Partner Institute for Computational Biology (PICB)

2010: DAAD-Dozent St. Luke’s Hospital, Quezon City (Manila) The Philippines

  2011-2013: DAAD-Dozent und Berater an der Pontificia Universidad Católica del Ecuador, Quito, Escuela de Ciencias Biológicas


European Molecular Biology Organization (EMBO)

American Genetics Society (GSA)

American Cell Biology Society (ASCB)

Gesellschaft für Genetik

Other Activities

Managing Editor of Chromosoma (1982-2001)

Member of the Editorial Board of Cell Research (Academia Sinica, Shanghai/China)

Member of the Management Group of the European Biochemistry Node for Interactions with China (EBNIC) of the European Union 1998-2001

Steering Committee of BIONET, an EU FP6 consortium on Bioethics (2006-2009)