1960-1963: Study of Biology/Chemistry at the Free University Berlin/Germany
1963-1967: Study of Biology/Genetics/Biochemistry at the Eberhard-Karls-University Tübingen/Germany
1967: Ph.D. at the Max-Planck-Institut für Biologie, Abt. Beermann, Tübingen/Germany
1967-1972: Research Associate, Max-Planck-Institut für Biologie, Abt. Beermann, Tübingen/Germany
1968: Postdoctoral Fellow, University of Edinburgh, MRC Group on the Mammalian Genome, Edinburgh/Scotland with P.M.B. Walker
1972: Habilitation “Biology”, Eberhard-Karls-University, Tübingen/Germany
1973: Habilitation “Molecular Biology”, University of Zürich/Switzerland
1973-1974: Oberassistent and Privatdozent, Institute of Molecular Biology II, University of Zürich/Switzerland
1974-1977: Head of an Independent Research Group at the Friedrich-Miescher-Laboratorium of the Max-Planck-Gesellschaft, Tübingen
1978-1997: Professor of Genetics, Head of Department of Genetics, Katholieke Universiteit Nijmegen/The Netherlands (Emeritus since 1997)
since 1985: Teaching and Research in the Guest Laboratory of the Max-Planck-Gesellschaft, Institute of Cell Biology, Academia Sinica,
since 1991 at the West China University of Medical Sciences, Chengdu/China
1991: Honorary Professor of the Academia Sinica, Institute of Cell Biology, Shanghai/China
1994: Honorary Professor of the West China University of Medical Sciences, Chengdu/China
1997- 2000: Guestprofessor, Faculty of Medicine, Johannes Gutenberg-University of Mainz/Germany
1998-2000: Member of the Managing Committee of the EBNIC (European Biotechnology Node for Interactions with China) an EU-funded activity of the EMBO (see: http://www.ebnic.org)
1998: Honorary Professor (Faculty of Biology), Johannes Gutenberg-University Mainz/Germany
2001: Honorary Professor Tongji Medical College Wuhan/China
2000-2008: Professor at the Shanghai Institutes for Biological Sciences (SIBS), Academia Sinica (long term lecturer of the German Academic Exchange Service (DAAD), and CAS-MPG Partner Institute for Computational Biology (PICB)
2010: DAAD-Dozent St. Luke’s Hospital, Quezon City (Manila) The Philippines
2011-2013: DAAD-Dozent und Berater an der Pontificia Universidad Católica del Ecuador, Quito, Escuela de Ciencias Biológicas
Memberships:
European Molecular Biology Organization (EMBO)
American Genetics Society (GSA)
American Cell Biology Society (ASCB)
Gesellschaft für Genetik
Other Activities
Managing Editor of Chromosoma (1982-2001)
Member of the Editorial Board of Cell Research (Academia Sinica, Shanghai/China)
Member of the Management Group of the European Biochemistry Node for Interactions with China (EBNIC) of the European Union 1998-2001
Steering Committee of BIONET, an EU FP6 consortium on Bioethics (2006-2009)